僥儞僌僇僗儀Dipturus tengu (Jordan and Fowler, 1903) (僈儞僊僄僀栚僈儞僊僄僀壢)丂暊柺幨恀丂
2014擭4寧偺嫑
僥儞僌僇僗儀Dipturus tengu (Jordan and Fowler, 1903) (僈儞僊僄僀栚僈儞僊僄僀壢)丂暊柺幨恀丂
丂僈儞僊僄僀栚乮Rajiformes丆偄傢備傞”僇僗儀”乯偼悽奅偵245庬埲忋偑抦傜傟, 慡悽奅偺挭壓懷偐傜戝棨幬柺堟傑偱峀偔暘晍偡傞掙惗惈擃崪嫑椶偱偡 (Ebert and Compagno, 2007)丏杮栚偼嵟傕斏塰偟偰偄傞擃崪嫑椶偱偁傝丆僄僀椶乮僄僀忋栚乯偺拞偱偼桞堦偺棏惗偱丆挿曽宍偺晅拝惈棏傪嶻偟, 旜晹偵偼旝庛側揹棳傪敪偡傞敪揹婍姱偑偁傝, 偦偟偰暊昲偑忋壓偵暘偐傟傞側偳偺摿挜傪傕偪傑偡 (McEachran and Dunn, 1998)丏僈儞僊僄僀栚偼僸僩僣價儗僇僗儀壢 (Arhynchobatidae), 儂僐僇僗儀壢 (Anacanthobatidae) , 偦偟偰僈儞僊僄僀壢 (Rajidae) 偺3壢偐傜峔惉偝傟傑偡 (Ebert and Compagno, 2007; 愇尨, 2012). 擔杮嶻僈儞僊僄僀壢嫑椶偼5懏12庬偑抦傟傜偰偍傝, 偦偺偆偪僥儞僌僄僀懏 Dipturus Rafinesque, 1810偵偼僈儞僊僄僀Dipturus sp., 僉僣僱僇僗儀D. macrocauda, 僝僂僇僗儀D. gigas, 偦偟偰杮庬偺4庬偑抦傜傟偰偄傑偡 (愇尨, 2012; 攇屗壀傎偐, 2013). 僥儞僌僄僀懏偼屌偔懢偄斅忬偺暙擃崪傪旛偊, 暙抂偺梼忬擃崪偑暙擃崪偲桙崌偡傞偙偲傗, 暙偑挿偔丆摢挿偵懳偡傞暙擃崪挿偑60 %埲忋偱偁傞偙偲, 惉弉僒僀僘偑慡挿 55 cm埲忋偲斾妑揑戝宆偱偁傞偙偲, 偍傛傃旜晹偺灆楍偑梇偱偼1楍, 帗偱偼3楍偱偁傞偙偲側偳偱懠懏偲嬫暿偝傟傑偡 (Ishiyama, 1958; Ishihara, 1987; 攇屗壀傎偐, 2013)丏Ishiyama (1958) 偼杮庬傪僞僀僾庬偲偟偰Tengujei 傪愝棫偟傑偟偨偑, 偙傟偼偺偪偵Dipturus 偺僔僲僯儉偱偁傞偲偝傟傑偟偨 (Stehmann, 1970).
丂僥儞僌僇僗儀偼媨忛導徏搰榩偱嵦廤偝傟偨帗屄懱傪傕偲偵, Jordan and Fowler (1903) 偵傛偭偰怴庬婰嵹偝傟傑偟偨. 杮庬偼撿僔僫奀偐傜杒奀摴慡堟偵偐偗偰偺悈怺45–400 m偵暘晍偟, 擔杮嶻僥儞僌僄僀懏偺側偐偱偼杒曽偐偮愺奀堟傑偱弌尰偡傞孹岦偺偁傞庬偱偡 (Ishiyama, 1967; 攇屗壀傎偐, 2013). 慡挿偼1 m傪挻偊, 僈儞僊僄僀栚嫑椶偺側偐偱偼拞宆偺庬偱偡. 杮庬偼”僥儞僌”偺柤偑帵偡傛偆偵, 暙偑挊偟偔怢挿偟, 偦偺愭抂偑愲傞偙偲傗, 旜晹偺敪揹婍偑偁傑傝敪払偣偢丆嵶挿偄旜傪桳偡傞偙偲, 偍傛傃暊昲偺暊柺偵暘晍偡傞姶妎岴偑斾妑揑柧椖偱偁傞偙偲側偳偱丆擔杮嶻偺杮懏懠庬偲嬫暿偱偒傑偡. 偟偐偟側偑傜, 暘椶宍幙偵朢偟偄杮栚嫑椶偱偼, 帗梇嵎傗惉挿偵傛傞曄堎偵壛偊偰, 庬偵傛偭偰偼挊偟偄屄懱曄堎傪帵偡偙偲偐傜, 庬偺摨掕偑崲擄側応崌傕偁傝傑偡.
丂僈儞僊僄僀栚嫑椶偺暘椶偵偼丆庬儗儀儖偺栤戣偺懠偵, 懏儗儀儖偵傕栤戣偑偁傝傑偡丏摿偵丆僥儞僌僄僀懏偼僆僇儊僄僀懏Okamejei Ishiyama, 1958偲偺椶帡惈偑巜揈偝傟偰偄傑偡丏斈悽奅揑側暘晍傪帵偡僥儞僌僄僀懏偵懳偟偰丆僆僇儊僄僀懏偼僀儞僪-惣晹懢暯梞偵屌桳偺僌儖乕僾偱, 擔杮偐傜偼僐儌儞僇僗儀O. kenojei 傗僣儅儕僇僗儀O. schmidti 側偳5庬偑抦傜傟偰偄傑偡. 僆僇儊僄僀懏偺摿挜偼懱挿偑彫偝偔, 偍偍傓偹慡挿40–50 cm掱搙偱惉弉偟 (僥儞僌僄僀懏偱偼堦晹偺庬傪彍偒慡挿1 m傪挻偊偰惉弉), 暙偼抁偔, 旜晹偺灆楍偼梇偱偼3楍丆帗偱偼5楍偲偝傟偰偄傑偡 (Ishiyama, 1958; Ishihara, 1987; 攇屗壀傎偐, 2013). 偟偐偟側偑傜, 椉懏偺拞娫揑側宍幙傪帵偡庬偑曬崘偝傟偰偍傝, 椺偊偽, D. oculus Last, 2008偼暙偑抁偔懱僒僀僘傕60 cm偵枮偨側偄側偳偺摿挜偱僆僇儊僄僀懏偵帡傑偡偑, 旜晹偺灆楍側偳偺摿挜偱偼僥儞僌僄僀懏偵堦抳偟傑偡. 擔杮嶻偺僥儞僌僄僀懏偺偆偪丆僈儞僊僄僀偼僥儞僌僄僀懏偲僆僇儊僄僀懏偺拞娫揑側摿挜傪傕偪傑偡. 僈儞僊僄僀偼懱僒僀僘偑斾妑揑彫偝偄偙偲傗, 暙傕斾妑揑抁偄偙偲側偳, 僆僇儊僄僀懏偵帡偨宍幙傪傕偪暪偣偰偄偨偨傔偵, 夁嫀偵偼僆僇儊僄僀懏偵暘椶偝傟偰偄偨偙偲傕偁傝傑偟偨 (Ishiyama, 1958; Ishiyama, 1967). 埲忋偺傛偆偵, 僥儞僌僄僀懏偲僆僇儊僄僀懏偺柧妋側嬫暿偼崲擄偱偁傝, 摿偵僥儞僌僄僀懏偼嵞掕媊偡傞昁梫偑偁傞偲偝傟偰偄傑偡 (McEachran and Fechhelm, 1982; Seret, 1986).
丂偝偰, 偙偺幨恀偺昗杮偵偮偄偰偱偡偑, 2014擭1寧24擔偺崅抦巗屼忯悾嫏峘偺戝庤孞傝栐嫏偺嫏楯偔偢偺拞偵崿偞偭偰偄傑偟偨. BSKU昗杮僐儗僋僔儑儞偵強憼偝傟偰偄傞僥儞僌僇僗儀偼慡晹偱13屄懱偱, 偦偺傎偲傫偳偑1950擭戙偵嵦廤偝傟偨傕偺偱, 偙偺昗杮偼5擭傇傝偺捛壛屄懱偵側傝傑偟偨. 杮庬傪娷傓僥儞僌僄僀懏偼屼忯悾嫏峘偺戝庤孞傝栐嫏偱偟偽偟偽崿妉偝傟偰偄傞傛偆偱偡丏偟偐偟, 懱僒僀僘偑戝偒偄偙偲偵壛偊, 崅抦偱偼巗応壙抣偑側偄杮懏嫑椶偼, 傎偲傫偳偺応崌慏忋偱攑婞偝傟偰偟傑偄傑偡. 堦曽, 僈儞僊僄僀椶偼搶杒偍傛傃杒奀摴偱偼怘梡嫑偲偟偰偺壙抣偑崅偔, 廐揷導偺搚嶈峘嵳傝偵偼“僇僗儀嵳”傝偲偄偆暿柤偑偁傞傎偳偱偡. 娯崙偱傕尵傢偢偲抦傟偨億僺儏儔乕側嫑偱偡. 傢偨偟傕惀旕僥儞僌僄僀懏嫑椶傪徿枴偟偰傒偨偄傕偺偱偡偑, 戝曄婱廳側僒儞僾儖偱偁傞偨傔, 嵦傟偨帪偵偼柪傢偢昗杮偵偡傞偙偲偱偟傚偆.
嶲峫暥專
Ebert, D. A . and L. J. V. Compagno. 2007. Biodiversity and systematics of skates (Chondrichthyes: Rajiformes: Rajoidei). Environ. Biol. Fish., (2007) 80: 111–124.
攇屗壀惔曯丒嶳揷攡朏丒棔郪惓岹丒嶳岥撝巕丒桍壓捈屓. 2013. 僈儞僊僄僀壢. 拞朧揙師 (曇), pp. 205–216, 1771–1773. 擔杮嶻嫑椶専嶕 慡庬偺摨掕. 戞3斉. 搶奀戝妛弌斉夛, 恅栰.
Ishihara, H. 1987. Revision of the western North Pacific species of the genus Raja. Japan. J. Ichthyol., 34 (3): 241–285.
愇尨丂尦. 2012. 僈儞僊僄僀栚嫑椶偺懏偺昗弨榓柤偺惍棟偲杒懢暯梞嶻慡庬偺昗弨榓柤儕僗僩. 斅橐椶尋媶夛曬, 48: 6–15.
Ishiyama, R. 1958. Studies on the rajid fishes (Rajidae) found in the waters around Japan. J. Shimonoseki Coll. Fish., 7 (2–3): 191–394, pls.1–3.
Ishiyama, R. 1967. Fauna Japonica. Rajidae (Piscea). Biogeographical Society of Japan, Tokyo. 鶨+84 pp., 32 pls.
Jordan, D. S. and H. W. Fowler. 1903. A review of the elasmobranchiate fishes of Japan. Proc. U.S. Natn. Mus., 26(1324): 593-674.
Last, P. R. 2008. New short-snout members of the skate genus Dipturus (Rajoidei: Rajidae) from Australian seas. Pages 53–98 in P. R. Last, W. T. White, J. J. Pogonoski and D. C. Gledhill, eds. Descriptions of new Australian skates (Batoidea: Rajoidei). CSRIO Marine and Atmospheric Research Paper No. 021, Hobert.
Last, P. R. and G. K. Yearsley. 2002. Zoogeography and relationships of Australasian skates (Chondrichthyes: Rajidae). J. Biogeogr, 29: 1627–1641.
McEachran, J. D. and K. A. Dunn. 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes; Rajidae). Copeia, 1998 (2): 271–290.
McEachran, J. D. and J. D. Fechhelm. 1982. A new species of skate from the western Indian Ocean, with comments on the status of Raja (Okamejei) (Elasmobranchii: Rajiformes). Proc. Biol. Soc. Washington, 95 (3): 440–450.
Seret, B. 1986. Deep water skates from Madagascar (Chondrichthyes, Rajoidei). Pages 256–260 in T. Uyeno, R. Arai, T. Taniuchi, and K. Matsuura, eds. Indo-Pacific fish biology. Ichthyological Society of Japan, Tokyo. 998 pp.
Stehmann, M. 1970. Vergleichend morphologische und anatomische Untersuchungen zur Neuordnung der Systematik der nordostatlantischen Rajidae (Chondrichthyes, Batoidei). Arch. Fischwiss., 21 (2): 73-164.
幨恀昗杮僨乕僞: BSKU 112349, 356 mm TL, 亰, 2014擭1寧24擔, 崅抦巗屼忯悾嫑巗応戝庤孞傝栐嫏 (嫽捗壂, 悈怺280– 320 m), 嵦廤仌幨恀: 嶰郪丂椛.
乮嶰郪丂椛乯
(C) BSKU Laboratory of Marine Biology, Kochi University
